The big thing to keep in mind is this: not all lead is the same. There is organic lead compounds (like the ones found in lead paint, leaded gasoline, etc) and inorganic lead (the metal stuff that is in bullets).
Organic lead is very easily absorbed into the bloodstream, either through ingestion or through skin contact. Lead acetate is a classic example of this. The Romans used to use it as a sweetener. Nowadays people who use the "dip" method to clean their rimfire suppressors are creating it. It is very easily absorbed through the skin and can be very harmful.
Inorganic lead, on the other hand, is not easily absorbed. That is why many have issues with the studies that are published. They don't ever say exactly what types of lead they are exposing their test subjects to, or what types of lead their subjects are subjected to on a normal basis.
In every link you posted, the discussion was around bullet fragments in meat. Every one of them points to studies that were conducted regarding the lead levels in people that came from other sources of exposure (such as lead paint). There has not, to my knowledge, been a definitive study on hunters over the course of their lives in regards to how the lead levels in their bloodstream increase over time as they eat game killed with lead bullets.
One of the best studies I have found, where they actually published their methods and findings is this one:
https://soarraptors.org/wp-content/uploads/NorthDakotaCDCreport.pdf
Of course, even when searching for it, and on the ND DNR website, the item that is noted most is: "The study shows a link between eating wild game shot with lead bullets and higher blood lead levels." While factually true, it is a bit misleading. What the study found (if you don't want to read the whole thing) is that people who ate wild game meat had a mean lead level of 1.27 μg/dL (microgram per deciliter). Those who did not eat wild game meat had a level of 0.84 μg/dL.
That sounds significant, right? It does until you read this statement from the study: "and none exceeded the Centers for Disease Control and Prevention’s level of concern of 10 ug/dl." Just for comparison, the current blood lead reference level recognized by the Centers for Disease Control and Prevention is 3.5 micrograms per deciliter (μg/dL).
So, those people studied in ND had lead levels less than half of the reference level of the CDC.
In regards to the primer question, the lead compound used in primers is of the organic kind. When you fire, it becomes vaporized and can be breathed in. That is where the danger lies. That is also one of the biggest factors when shooting in indoor ranges. So, if you are not using lead projectiles but still shooting in indoor ranges, you are actually at greater risk for harmful lead exposure. It is also one of my peeves with these studies. I have yet to find one where one of the survey questions is "how much, on average do you shoot, and is it indoors or outdoors". Without that data, we cannot rule out that the increased in lead concentration in the blood is from just shooting a lot rather than eating meat with lead fragments in it.
And before anyone asks or implies, I spent quite a bit of time in my organic and inorganic chemistry classes in school learning the difference between the two types of compounds. They are not the same.