I will add my $0.02 here.
According to the article a lock on a gate was either cut or shot off. If that lock was on a gate that separated public land from private and the hunter was the one who removed the lock, then he was absolutely trespassing. Also, if you look at the map provided, the landlocked portions of public are very small in comparison and are not even close to the boundaries. This leads me to two conclusions. First, these are not elk that are hanging out exclusively on public land (like the ones in the corner crossing case). They spend the majority of their time on the private land that is owned and managed by the landowner. This most likely isn't a case of "the landowner is treating this public land herd as their own" like some have postulated here. Second, the landlocked parcels of public are pretty deep into the property. According to the article, the hunter had to travel "a mile" into the property to get to the public. Again this isn't just a "I accidentally crossed a boarder" or a corner-crossing thing. This is willful trespassing (provided it happened as described in the article).
As for the landowners treating the animals as "theirs". Myself and a couple of my best friends own several contiguous tracts of land in WV. All told it is a pretty large parcel for an Eastern state (837 acres). Two years ago, a large tract of paper company land that is along our SE border was leased to the state for use as state game lands. Since that time we have had a serious influx of people traveling down the private road that leads to our camp, and in more than one instance we have had people cross onto our property either hunting or trailing deer/bears. While I don't believe that we "own" the animals, I do believe that due to the price that we paid for the land, the money that we spend every year on taxes for that land, seed, fuel, maintenance of equipment, feed, general upkeep of the property, etc. entitles us to have exclusive rights to hunt those animals while they are on our property as well as entitling us to access that same public land in an area that would require others to have to travel a significant distance in order to get to from the main access points of the public land. I also don't agree that we should be forced to allow access to that public land through our land (there are a couple of places where it is only a couple hundred yards from the road to the public boundary). There are other access points and honestly, ours was private long before that piece was public (we were their first).